Publications

Hughes N.C. and Rushton A.W.A. 1990. Computer-aided reconstruction of a late Cambrian ceratopygid trilobite from Wales and its phylogenetic implications. Palaeontology  33:429-445.

Jell P.A., Hughes N.C. and Brown A.V. 1991. Late Cambrian (post-Idamean) trilobites from the Higgins Creek area, western Tasmania. Memoirs of the Queensland Museum 30:455-485.

Hughes N.C. 1991. Morphological plasticity and genetic flexibility in a Cambrian trilobite. Geology 19:913-916.

Hughes N.C. and Droser M.L. 1992. Trace fossils from the Phe Formation (Lower Cambrian), Zanskar valley, northwestern India. Memoirs of the Queensland Museum 32:139-144.

Hughes N.C. and Jell P.A. 1992. A statistical/computer graphic technique for assessing variation in tectonically deformed fossils and its application to Cambrian trilobites from Kashmir. Lethaia 25:317-330.

Hughes N.C. 1993. Distribution, taphonomy, and functional morphology of the Upper Cambrian trilobite Dikelocephalus. Milwaukee Public Museum Contributions in Biology and Geology 84:1-49.

Labandeira C.C. and Hughes N.C. 1994. Biometry of the Late Cambrian trilobite Dikelocephalus, and its implications for trilobite systematics. Journal of Paleontology 68:492-517.

Hughes N.C. 1994. Ontogeny, intraspecific variation, and systematics of the Late Cambrian trilobite Dikelocephalus. Smithsonian Contributions to Paleobiology 79:1-89.

Droser M.L., Hughes N.C. and Jell P.A. 1994. Palaeoecology of Cambro-Ordovician nearshore sandstones: trace fossil evidence from Mootwingee, New South Wales, Australia. Lethaia 27:273-283.

Hughes N.C. 1995. Trilobite taphonomy and taxonomy: a problem and some implications. Palaios 10:283-285.

Hughes N.C. and Labandeira, C.C. 1995. The stability of species in taxonomy. Paleobiology 21:401-403.

Hughes N.C. and Chapman R.E. 1995. Growth and variation in the Silurian trilobite Aulacopleura konincki and its implications for trilobite palaeobiology. Lethaia 28:333-353.

Rushton A.W.A and Hughes N.C. 1996. Biometry, systematics, and biogeography of the late Cambrian trilobite Maladioidella abdita. Transactions of the Royal Society of Edinburgh: Earth Sciences 86:247-256.

Hughes N.C., Gunderson G.O. and Weedon M.J. 1997. Circumocular suture and visual surface of "Cedaria" woosteri  (Trilobita, Late Cambrian) from the Eau Claire Formation, Wisconsin. Journal of Paleontology 71:103-107.

Hughes N.C. and Hesselbo S.P. 1997. Stratigraphy and sedimentology of the St. Lawrence Formation, Upper Cambrian of the northern Mississippi Valley. Milwaukee Public Museum Contributions in Biology and Geology 91:1-50.

Jell P.A. and Hughes N.C. 1997. Himalayan Cambrian trilobites. Special Papers in Palaeontology 58:1-113.

Hughes N.C. 1997. Cambrian trilobite biostratigraphy for the Himalaya. Current Science 73:736-737.

Fortey R.A. and Hughes N.C. 1998. Brood pouches in trilobites. Journal of Paleontology 74:638-649.

Hughes N.C. and Jell P.A. 1999. The biostratigraphy and biogeography of Himalayan Cambrian trilobites. in Macfarlane A, Sorkhabi R.B. and Quade J. (eds). Himalaya and Tibet: Mountain Roots to Mountain Tops, Boulder Colorado. Geological Society of America Special Papers 328:109-116.

Webster M. and Hughes N.C. 1999. Compaction-related deformation in well-preserved Cambrian olenelloid trilobites and its implications for fossil morphometry. Journal of Paleontology 73:355-371.

Hughes N.C. and Cooper D.L. 1999. Paleobiologic and taphonomic aspects of the "granulosa" trilobite cluster, Kope Formation (Upper Ordovician, Cincinnati region). Journal of Paleontology 73:306-319.

Hughes N.C.1999. Statistical and imaging methods applied to deformed fossils. pp. 127-155. in Harper, D.A.T. (ed.). Numerical Palaeobiology: computer-based modelling and analysis of fossils and their distributions. John Wiley Press, London. 468 pp.

Gaines R.R., Droser M.L. and Hughes N.C. 1999. The ichnological record in Ordovician mudstones: examples from the Cincinnatian strata of Ohio and Kentucky (USA). Acta Universitatis Carolinae - Geologica 43:163-165.

Hughes N.C., Chapman R.E. and Adrain J.M.1999. The stability of thoracic segmentation in trilobites: a case study in developmental and ecological constraints. Evolution and Development 1:24-35.

Hughes N.C., Gunderson G.O. and Weedon M.J. 2000. Late Cambrian conulariids from Wisconsin and Minnesota. Journal of Paleontology 74: 828-838.

Hughes N.C. 2000. Ecological evolution of Cambrian trilobites. p. 370-403. in  Zhuravlev A.Yu. and Riding R. (eds.). The Ecology of the Cambrian Radiation Columbia University Press, New York. 525 pp.

Hughes N.C., Collier F.J., Klussendorf J., Lipps J.H., Taylor W.L., and White R.D. 2000. Fossil invertebrate and micro-fossil collections: Kinds, uses and users. in White, R.D. and Allmon, W.D.(eds.). Guidelines for the Management and Curation of Invertebrate Fossil Collections. Paleontological Society Special Publications 10: 25-35.

Webster, M., Sheets, H.D., and Hughes N.C. 2001. Allometric patterning in trilobite ontogeny: Testing for allometric heterochrony in Nephrolenellus. pp 105-144. in Zelditch, M.L. ed. Beyond Heterochrony. Wiley-Liss, New York. 371 pp.

Hughes N.C. and Chapman R.E. 2001. Morphometry and phylogeny in the resolution of paleobiological problems - unlocking the evolutionary significance of an assemblage of Silurian trilobites. p. 29-54. in Adrain, J.M., Edgecombe, G.E. and Lieberman, B.S. eds. Fossils, Phylogeny, and Form: An Analytical Approach. Kluwer Academic/Plenum Press. New York. 402 pp.

Hughes N.C., Peng Shanchi and Luo Huilin 2002. Kunmingaspis (Trilobita) putatively from the Yunling Collage, and the Cambrian faunal history of the eastern Himalayan syntaxial region. Journal of Paleontology 76: 709-717.

Hughes N.C. 2002. Late Middle Cambrian trace fossils from the Lejopyge armata horizon, Zanskar Valley, India, and the use of Precambrian/Cambrian ichnostratigraphy in the Indian subcontinent. Special Papers in Palaeontology 67:135-151.

Hughes N.C. 2003. Trilobite body patterning and the evolution of arthropod tagmosis. BioEssays 25:386-395.

Myrow P.M., Hughes N.C., Paulsen T.S., Williams I.S., Parcha S.K., Thompson K.R., Bowring S.A., Peng S.-C., and Ahluwalia A.D. 2003. Integrated tectonostratigraphic reconstruction of the Himalaya and implications for its tectonic reconstruction. Earth and Planetary Science Letters 212:433-441.

Minelli A., Fusco G. and Hughes N.C. 2003. Tagmata and segment specification in trilobites. Special Papers in Palaeontology 70:31-43.

Hughes N.C. 2003. Trilobite tagmosis and body patterning from morphological and developmental perspectives. Integrative and Comparative Biology 43:185-206.

Peng Shanchi, Babcock L.E., Hughes N.C. and Lin Huanling. 2003. Cambrian shumardioids from northwestern Hunan, China. Special Papers in Palaeontology 70: 197-212.

Fusco, G. Hughes, N.C, Webster, M, and Minelli, A. 2004. Exploring developmental modes in a fossil arthropod: growth and trunk segmentation of the trilobite Aulacopleura konincki. American Naturalist 163:167-183.

Hughes, N.C. and Heim, N.A. 2005. Cambrian: chronology, environments, and evolution. in Selley R.C, Cocks L.R.M. and Plimer I.R. Encyclopedia of Geology. Elsevier Press, Oxford, Vol. 4, pp. 163-174.

Hughes N.C., Peng Shanchi, Bhargava O.N., Ahluwalia A.D., Walia S., Myrow P.M. and Parcha S.K. 2005. Early Tsanglangpuan (late early Cambrian) trilobites from the Nigali Dhar syncline and the Cambrian biostratigraphy of the Tal Group, Lesser Himalaya, India. Geological Magazine 142: 57-80.

Hughes, N.C., Myrow P.M., Peng Shanchi, and Parcha S.K. 2005. Establishing a chronostratigraphic and paleoenvironmental framework for the Cambrian of the Himalaya and its geological implications. Acta Micropalaeontologica Sinica 22 Supplement: 68-70.

Jacobs D.K., Hughes N.C., Fitz-Gibbon S.T, and Winchell C.J. 2005. Terminal addition, the Cambrian radiation and the Phanerozoic evolution of bilaterian form. Evolution and Development 7: 498-514.

Simpson A., Hughes N.C., Kopaska-Merkel D.C. and Ludvigsen, R. 2005. Development of the caudal exoskeleton of the pliomerid trilobite Hintzeia plicamarginis new species. Evolution and Development 7: 528-541.

Hughes, N.C. 2005. Trilobite construction: building a bridge across the micro- and macroevolutionary divide. p. 139-158. in Briggs, D.E.G. (ed.) Evolving Form and Function: Fossils and Development: Proceedings of a symposium honoring Adolph Seilacher for his contributions to paleontology, in celebration of his 80th birthday. Peabody Museum of Natural History, Yale University. New Haven. 288pp.

Myrow P.M., Thompson K.R., Hughes N.C., Paulsen T.S., Sell B.K., and Parcha S.K. 2006. Cambrian stratigraphy and depositional history of the northern Indian Himalaya, Spiti Valley, north-central India. Geological Society of America Bulletin 118:491-510.

Myrow P.M., Snell K.E., Hughes N.C., Paulsen T.S., Heim N.A., and Parcha S.K. 2006. Cambrian depositional history of the Zanskar Valley region of the Indian Himalaya: Tectonic implications. Journal of Sedimentary Research 76:364-381.

Hunda B.R., Hughes N.C. and Flessa K. 2006. Trilobite taphonomy and temporal resolution in the Mt. Orab Shale (Upper Ordovician, Ohio, U.S.A.). Palaios 21:26-45.

Hughes N.C., Minelli A., and Fusco G. 2006. The ontogeny of trilobite segmentation: a comparative approach. Paleobiology 32:603-628.

Hughes, N.C. 2007. The Evolution of Trilobite Body Patterning. Annual Reviews in Earth and Planetary Sciences 35:401-434.

Hunda B.R and Hughes N.C. 2007. Evaluating paedomorphic heterochrony in trilobites: the case of the diminutive trilobite Flexicalymene retrorsa minuens from the Cincinnatian Series (Upper Ordovician), Cincinnati region. Evolution and Development 9:483-496.

Paulsen T.S., Demosthenous C.M., Myrow P.M., Hughes N.C., and Parcha S.K. 2007. Paleostrain stratigraphic analysis of calcite twins across the Cambrian-Ordovician unconformity in the Tethyan Himalaya, Spiti and Zanskar valley regions, India. Journal of Asian Earth Sciences 31:44-54.

Hughes, N.C. 2007. Strength in numbers: high phenotypic variance in early Cambrian trilobites and its evolutionary implications. BioEssays 29:1081-1084.

Zhu X.-J., Hughes N.C., and Peng S.-C. 2007. On a new species of Shergoldia Zhang & Jell 1987 (Trilobita), the family Tsinaniidae, and the Order Asaphida. Memoirs of the Australasian Association of Palaeontologists 34:243-253.

Hughes N.C., Haug J.T. and Waloszek D. 2008. Basal euarthropod development: a fossil-based perspective. p. 279-296. in Minelli A. and Fusco G. eds. Evolutionary Pathways - Key Themes in Evolutionary Developmental Biology. Cambridge University Press. 426pp.

Peng S.-C., Yang X.-F. and Hughes N.C. 2008. The oldest stalk-eyed trilobite, Parablackwelderia Kobayashi, 1942 (Damesellinae, Cambrian), and its occurrence in Shandong, China. Journal of Paleontology 82:842-850.

Webster M., Gaines R.R. and Hughes, N.C. 2008. Microstratigraphy, trilobite biostratinomy, and depositional environment of the "lower Cambrian" Ruin Wash Lagerstätte, Pioche Formation, Nevada. Palaeogeography, Palaeoclimatology, Palaeoecology 264:100-122

Balint M. and Hughes, N.C. 2008. Segment variation in a Cambrian trilobite in the context of the Burgess Shale fauna. In Rábano, I., Gozalo, R. and Garcia-Bellido, D. (eds.). Advances in Trilobite Research. Cuadernos del Museo Geominero, No. 9, Instituto Geológico y Minero de España, Madrid, 29-32.

Hong P.S. Hughes N.C. Fusco G. and Zhang X.G. 2008. Trunk segmentation of the early Cambrian eodiscid trilobite Neocobboldia chinlinica. In Rábano, I., Gozalo, R. and Garcia-Bellido, D. (eds.). Advances in Trilobite Research. Cuadernos del Museo Geominero, No. 9, Instituto Geológico y Minero de España, Madrid, 181-184.

Hughes N.C. and Fusco G. 2008. Growth progression in trilobites: a preliminary study. In Rábano, I., Gozalo, R. and Garcia-Bellido, D. (eds.). Advances in Trilobite Research. Cuadernos del Museo Geominero, No. 9, Instituto Geológico y Minero de España, Madrid, 195-198.

Hughes N.C. Peng S. and Parcha S.K. 2008. Cambrian trilobites from the Parahio and Zanskar Valleys, Indian Himalaya: a synopsis. In Rábano, I., Gozalo, R. and Garcia-Bellido, D. (eds.). Advances in Trilobite Research. Cuadernos del Museo Geominero, No. 9, Instituto Geológico y Minero de España, Madrid, 199-200.

Hunda B.R. Thompson A. Webber A.J. and Hughes N.C. 2008. Short-term paleoenvironmentally-related morphological variation in the Ordovician trilobite Flexicalymene granulosa. In Rábano, I., Gozalo, R. and Garcia-Bellido, D. (eds.). Advances in Trilobite Research. Cuadernos del Museo Geominero, No. 9, Instituto Geológico y Minero de España, Madrid, 201-204.

McKenzie N.R. Hughes N.C. Myrow P.M. Bhargava O.N. Tangri S.K and Ghalley K.S. 2008. Late Cambrian (Furongian) trilobites of the Pele La Group, Black Mountain region, Bhutan, and their significance for reconstruction of the early Paleozoic Gondwana margin. In Rábano, I., Gozalo, R. and Garcia-Bellido, D. (eds.). Advances in Trilobite Research. Cuadernos del Museo Geominero, No. 9, Instituto Geológico y Minero de España, Madrid, 265-268.

Paterson J.B. Hughes N.C. and Chatterton B.D.E. 2008. Trilobite clusters. What do they tell us? A preliminary investigation. In Rábano, I., Gozalo, R. and Garcia-Bellido, D. (eds.). Advances in Trilobite Research. Cuadernos del Museo Geominero, No. 9, Instituto Geológico y Minero de España, Madrid, 313-318.